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This fermentation process changes the flavour profile of the tobacco as there is less oxygen touching all parts of the tobacco because the flake slices are a part of a larger block of compressed tobacco

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References Bousquet J, Khaltaev N, Cruz AA, Denburg J, Fokkens WJ, Togias A, et al

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Cells were immunoprecipitated with Sp1 or SIRT1 antibodies or normal IgG (Cell Signaling Technology) and then amplified by PCR